CAPELIN

Mallotus villosus


Assessment report
Published by

Marine and Freshwater Research Institute, Iceland

Published

06 June 2025

Key signals

  • The size of the capelin stock and the catches have fluctuated between years and are reflecting the size of the incoming cohort at any given time. Since changes occurred in the stock’s nursery and feeding grounds since around 2000, both stock size and catches have generally been smaller.

  • The size of the fishable stock in the last two seasons was estimated small, and therefore no catches were allowed for the 2023/2024 season and very small catches in 2024/2025, with an advice of 8 589 tonnes.

  • The index of the abundance of juvenile capelin in the autumn survey of 2024 was 58.9 billion individuals, which according to the harvest control rule leads to an initial advice for the 2025/2026 season of 46 384 tonnes. This initial advice will be revised following the autumn survey in September 2025.

General information

The capelin is a small pelagic schooling fish. It is a cold-water species that inhabits arctic and subarctic waters in the North Atlantic and North Pacific. Capelin in the Iceland-East Greenland-Jan Mayen area is considered to be a single stock, and is denoted here after as the IEGJM capelin stock.

Fisheries of capelin began in Iceland in the mid-1960s. Besides being commercially important capelin is a key species in the marine ecosystem off Iceland. It feeds on small zooplankton, especially copepods but also amphipods and krill, but is itself an important food for cod, saithe, haddock, halibut and other commercial fish. It is also important prey for whales and birds. Thus, capelin is an important link in the transfer of energy and nutrients to the upper levels of the food web. Preferred temperature for capelin is usually 1-3°C during feeding migration and it is then often at the southern extent of cold Arctic water. When the adult capelin migrates from the feeding areas far north of Iceland, east and southeast of Greenland, it is mainly considered to approach the shelf break north of Iceland and then migrate clockwise around the island. Part of the main migration continues all the way towards the coast west of Iceland. Spawning takes place in shallow water in March-April in relatively warm seas south and southwest coast of the country. Spawning has also been observed in coastal areas north of Iceland but the extent of that spawning has been considered small compared to the number that spawns in the south. The majority of capelin dies after spawning, usually at the age of 3 years, although mostly females may survive spawning (Christiansen et al., 2008). The migrations of capelin contribute to a huge transfer of energy into the ecosystem of the Icelandic continental shelf. Larvae and juveniles drift clockwise along the continental shelf north and east of Iceland and in variable quantities towards Denmark Strait and to the shelf of East Greenland. Nursery grounds of capelin are in the waters north of Iceland and increasingly on the continental shelf of East Greenland (Bardarson et al., 2021) since the early 2000s.

Fisheries

No fisheries on the IEGJM capelin stock took place in the 2023/2024 fishing season. A total of 9 439 t catches were fished during the winter of 2025 (Figure 1). Total historical catch of the stock is shown in Figure 1 by season and distribution of the catches of the Icelandic fishing fleet by year can be seen on Figure 2.

Figure 1: Capelin. Total catch (in thousand tonnes) of IEGJM capelin since 1963/64 by season.
Figure 2: Capelin. Distribution of catches by the Icelandic fishing fleet based on logbooks.

Changes in fishing technology and fishing patterns

A total of 9 439 t catches were fished during the winter of 2024/2025 fishing season primarily by purse seine. Historically, most of the catches have been taken in purse seine but a variable amount has been taken with pelagic trawl through the fishing seasons, related to the size of the TAC and when it is issued. Discards have been considered negligible.

Stock assessment

Capelin acoustic assessment surveys

The IEGJM capelin stock has been assessed by acoustics annually since 1978. The surveys have been conducted in autumn (September-December) and in winter (January-February). An overview is given in the Northwestern Working Group NWWG, ICES stock annexe.

Autumn survey 2025

The Autumn survey was conducted with the aim of assessing both the immature and the maturing part of the stock (Bardarson et al., 2024). Since 2010, the autumn surveys have started in September (and partly in late August), a month earlier than in the preceding years because of difficulties in covering the stock due to drift ice and weather during later months at the more north-westerly feeding grounds of the stock since early 2000s (Vilhjálmsson, 2007).

The survey was conducted on the behalf of MFRI by the r/v Arni Fridriksson and r/v Tarajoq on behalf of GINR (Greenland Institute of Natural Resources) (Figure 3). The survey area was on and along the shelf edge off East Greenland from about 64°30´N towards about 72°15´N, also covering the Denmark Strait and the slope off northwest Iceland. The Iceland Sea and Kolbeinsey ridge were only briefly scouted due to time constraints and for same reason hydrographic measurements and zooplankton sampling were limited due to bad weather conditions. There were also considerable delays due to stormy weather.

In general, drift ice only limited the coverage of the survey vessels in a few regions although icebergs and a lack of information on bottom topography occasionally affected routes and limited the extent of transects towards the Greenlandic coast (Figure 3).

Maturing capelin was mainly observed outside the Icelandic continental shelf North and Northwest of Iceland (Figure 4). In the western Denmark Strait maturing capelin was mixed with immature capelin, while mainly maturing capelin was found further west. In contrast to last year, the distribution of mature capelin reached only a relatively short distance east of Denmark Strait and that is again a drastic change from observations in autumn 2022, when mature capelin was found on or along the East Greenland continental shelf south, east and north of Scoresby Sound. Further, no capelin was found near the western part of the Jan Mayen ridge or at the Kolbeinsey ridge. In general, there were no signs of significant quantities of capelin east of Kolbeinsey ridge nor along Icelandic shelf edges. Juveniles (0-group) of various species, including capelin (although not quantified) were observed along the continental shelf north and northwest of Iceland and along the southwestern coverage of the Greenlandic shelf. Immature capelin was found along the Greenlandic shelf, dominating in southwestern part of the survey area and Denmark Strait (Figure 4).

Figure 3: Capelin. The relative density of acoustic backscatter as NASC during 21 August–1 October 2024. Survey tracks by r/v Arni Fridriksson, f/v Polar Ammassak and r/v Tarajoq are shown in different colors.
Figure 4: Capelin. Transects of the participating vessels and proportion mature within capelin samples and regional allocation in assessment during the autumn survey in 21 August–1 October 2024.

The total biomass estimate was 624 722 tonnes. Table 1 gives information on the age-disaggregated numbers and biomass of the capelin stock components.

Table 1: Capelin. Estimated stock size of IEGJM capelin total stock in numbers (N) and biomass (tonnes) by age (years) and length (cm) from the autumn acoustic survey (21 August to 1 October 2024, see Figure 3).
Length age1 age2 age3 age4 Total Number Total Biomass (tonnes)
8.5 732 0 0 0 732 1551
9 2506 0 0 0 2506 6651
9.5 6544 0 0 0 6544 21177
10 10263 0 0 0 10263 38412
10.5 10947 0 0 0 10947 47360
11 7424 0 0 0 7424 36682
11.5 5555 0 0 0 5555 32184
12 5990 0 0 0 5990 39622
12.5 2834 138 0 0 2972 22260
13 2203 130 0 0 2334 19963
13.5 1381 310 0 0 1691 16254
14 891 785 49 0 1725 18679
14.5 457 1043 89 0 1589 20307
15 0 1727 0 0 1727 24940
15.5 49 1678 49 0 1776 28316
16 0 1513 81 0 1594 28871
16.5 0 2269 81 0 2350 46868
17 0 1955 191 0 2146 47007
17.5 0 1805 127 130 2063 50780
18 0 1107 127 32 1266 33371
18.5 0 640 64 64 768 22299
19 0 255 49 32 336 10505
19.5 0 127 64 99 290 8857
20 0 0 49 0 49 1809

Historical trend of mean weight of immature 1 to 3 year old capelin during the autumn survey is shown in Figure 5, as well as the trend of the mean weight of the mature capelin 1 to 4 year old capelin in Figure 6. The mean weight of 1 and 2 year old immature capelin has increased over time for all age classes, especially since 2000. Older age groups consistently show higher mean weights, with 3 year old immature capelin (Figure 5) and 4 year old capelin (Figure 6) exhibiting the most variability in mean weight.

Figure 5: Capelin. Mean weight of 1 to 3 year old immature capelin in autumn surveys since 1978. Colors are indicative of the different age classes and the lines represent the average weight for each class from 1978 to 2024.
Figure 6: Capelin. Mean weight of 1 to 4 year old mature capelin in autumn surveys since 1978.Colors are indicative of the different age classes and the lines represent the average weight for each class from 1978 to 2024.

Predation model results and resulting final advice 2024/2025

Following the completion of surveys, estimations of stock parameters and their uncertainty 100 thousand bootstrap replicates of SSB were used as starting values for predation model runs. Results from the predation model runs are given in Table 2 and shown in Figure 8 and Figure 9.

Table 2: Capelin. Quantiles and mean of SSB at time of spawning (15. March) and total predator consumption in thous. tonnes based on the predation model
mean 5% 25% 50% 75% 95%
SSB 210.19 113.33 162.11 203.34 250.67 330.94
Predation 114.81 72.98 94.67 112.25 132.40 165.10

The model (Ices 2023, ICES 2024) is designed to cover predation on the main spawning migration of capelin. A schematic description of the model is in Figure 7.

Figure 7: Capelin. Top: The 3 regions used in the simulations of predation on the capelin migration starting in the east, yellow arrows show the migration route. Bottom: Schematic showing proportional distribution of migrations and catches in the predation model as discretized over 2-week intervals. Catches are shown as fractions below horizontal arrows and clockwise migrations are given as percentages over vertical arrows.

The predation model (Ices 2023, ICES 2024) applies to the stock component that migrates the clockwise route around Iceland. In most years, majority of the stock has migrated that route and nearly all the catches have been taken from that component. All the capelin stock is assumed to be in the east on 15th of January, and on 15th of March, it is assumed that all the capelin stock spawns in the south and southwest, with a higher proportion in the southwest. The predators (cod, haddock and saithe) are assumed to be stationary during the period of capelin migration and their spatial distribution is obtained from the demersal survey in March from 1985 to the previous year. The total abundance of each predator is predicted for the current year based on assessment in the previous year.

Figure 8: Capelin. Summary of results from the 2024 autumn and winter 2025 acoustic survey and predation model. Biomass survey estimates of mature capelin (top-left), the projected spawning stock biomass left for spawning based on the predation model (top-right), predicted predation 15 January – 15 March (bottom-left) and the applied predation mortality (bottom-right).
Figure 9: Predicted development of the SSB with 8 589 catch based on the predation model. Blue line indicates Blim = 114 000 tonnes (Ices 2023, ICES 2024).
Table 3: Capelin. Assessment and advice for capelin in the IEGJM area for 2025. Fishable stock and CV the coefficient of variance of the survey estimates of the fish stock size. The advice is based on the predation model in the HCR.
Autumn January (?@fig-nasc6) February (?@fig-nasc9) Total Advice
Fishable stock (thousand tonnes) 318* 180 98.2 278.20 8589
CV 0.25 0.36 0.33 0.26 NA
Surveyed area East Greenland East of Kolbeinsey Western area NA NA

Initial advice 2025/2026

The initial advice for 2025/2026 is above the zero catch because the index for immature fish from the autumn acoustic survey in 2024 (see ?@sec-autumn2024) is 58.9 billion, and is above the Utrigger value (50 billion). Based on this rule (see Section 8.1), the initial TAC should be 46 384 t for the fishing season starting 15 October 2025 to 15 April 2026 (Figure 10).

Figure 10: Capelin. Catch advice according to the HCR for initial TAC, based on the relationship between final TAC and estimated number of immature capelin the previous autumn. The figure shows the estimated final TAC (black unbroken line) and the initial (preliminary) TAC (blue dashed line). The latter is set using a Utrigger (red vertical line) of 50 billion immature fish, with a cap on the initial (preliminary) TAC of 400 kt. The green lines show the index value from the autumn survey 2024, with the corresponding initial TAC for 2025/2026 shown on the y-axis.

Intermedia advice 2025/2026

The Intermedia advice for 2025/2026

Management

Agreed management plan and HCR

The Coastal States (Iceland, Greenland, and Norway) have agreed (Anon. 2015; Anon. 2023) to use the following harvest control rule as the basis for management, following the rule developed and revised by ICES (ICES, 2015; ICES, 2023).

The objective of the Harvest Control Rule is to set a final TAC which ensures, with a 95% probability, that a minimum of 114,000t (Blim) remains for spawning. This is achieved by a series of acoustic surveys from September - February and the TAC is determined in three stages; an initial TAC, an intermediate TAC and a final TAC.

  1. The initial TAC for the next fishing season is set based on estimates of the immature part of the stock following an acoustic survey in autumn (September - October).

    Two fixed points are defined:

    1. Utrigger = 50 billion immature capelin.

    2. TACMax = 400,000 tonnes for U > 127 billion immature capelin.

    The method for setting the initial/preliminary TAC is:

    • TAC = 0 if Uimm < 50 billion.

    • TAC = 5.2 x (Uimm - Utrigger) thous. tonnes for Uimm in the range 50–127 billion.

    • TAC = 400,000 tonnes if Uimm > 127 billion.

  2. The intermediate TAC for the current fishing season is set following the acoustic survey in autumn. Estimates on capelin spawning stock biomass (SSB) with uncertainty estimate are combined with data on predicted predator stocks size and distribution. The estimates are fed into a predation model run with varying catches until spawning in March. The intermediate TAC is set at 2/3 of the catch giving p(SSB < Blim = 114,000 tonnes) < 0.05.

  3. The final TAC is set following an acoustic survey in winter (January-February). Estimates on capelin spawning stock biomass with uncertainty estimate are combined with data on predicted predator stocks size and distribution. Final TAC is set based on all acoustic surveys on the mature part of the stock from autumn to winter. The estimates are fed into a predation model run with varying catches until spawning in March. The final TAC is set at the catch giving p(SSB < Blim = 114,000 tonnes) < 0.05.

Benchmark history

Since early 1980s the stock was managed according to an escapement strategy, leaving 400 000 tonnes for spawning (the uncertainty of the estimates was not considered). To predict the TAC for the next fishing season a model was developed in the early 1990s (Figure 11) (Gudmundsdottir and Vilhjálmsson, 2002). These models were not endorsed by the benchmark working group WKSHORT 2009.

A new HCR and management plan was endorsed by the benchmark working group WKICE in 2015 and WKCAPELIN in 2022 with minor adjustments and agreed upon by the coastal states (Figure 11). See WKICE (ICES, 2015), WKCAPELIN (ICES, 2023) and the Stock Annex for the capelin in the IEGJM area.

During WKICE, a Blim of 150 000 tonnes was defined (ICES, 2015) and following WKCAPELIN Blim was changed to 114 000 tonnes (Figure 11).

The initial (preliminary) quota follows a simple forecast that is based on a linear relation between historic observations of the abundance of juveniles from the acoustic autumn surveys and the corresponding final TACs nearly 1½ year later. Based on this rule, the advice on the initial quota for an upcoming fishing season is given.

The intermediate and final TACs are set so that there is at least 95% probability that there will be at least 114 000 tonnes (= Blim ) of mature capelin left for spawning at the spawning time (15 March). This was done for the first time in 2015/2016 by the Icelandic Marine Research Institute.

WKCAPELIN advised that in addition to the agreed management plan, the intermediate TAC following the autumn survey be set at 2/3 of the results of the predation model.

Figure 11: Capelin. Initial TAC, agreed final TAC, and catch as reported to ICES.

Reference points

Blim was set at 114 000 tonnes by WKCAPELIN (ICES, 2023). No other reference points are defined for this stock.

State of the stock

The fishable stock was estimated to 318 000 tonnes in October 2024 and 278 000 tonnes in January and February 2025. The predation model (ICES, 2015), accounting for catches (in this case winter catch of 8 589 t) and predation between surveys and spawning by cod, saithe and haddock, estimated SSB to 180 710 tonnes which were left for spawning in spring 2025 (Table 2). Given the uncertainty estimates, there was 95% probability that at least 114 000 tonnes was left for spawning, if the catches taken amounted to 8 589 tonnes, which represents the final TAC for capelin in the fishing year 2024/2025. The acoustic estimate of immature capelin from the autumn survey in October 2024 was 58.5 billion. Initial advice of 46 384 t for the fishing season 2025/26 on the basis of that amount is issued by ICES and MFRI in June 2026.

Figure 12: Capelin. Index of immature capelin from acoustic surveys in autumn since 1981.

The estimated SSB at spawning time (March-April) has been recompiled for 1981-2023 (Figure 13), using the model adopted in 2015 and 2023, i.e. taking into account uncertainty in the acoustic measurements and using the predation model adopted in 2015. Uncertainty in acoustic measurements was recompiled for the years 2002-2006 and 2012-2014 by recalculating the acoustic indices and bootstrapping the results. Additionally, uncertainty was available for the years since 2015 when the advice was given based on the new HCR. For earlier years, the CV in the acoustic measurements was estimated by looking at survey reports as well as text from (Vilhjálmsson, 1994). The estimated CV was in the range of 0.15-0.25 and was included as a lognormal multiplier on available average values from the same sources.

Figure 13: Capelin. Median, 5th and 95th percentile of SSB distribution at spawning time (March–April) since 1981. Blim = 114 kt.

Uncertainties in assessment and forecast

The uncertainty of the assessment and forecast depends largely on the quality of the acoustic surveys in terms of coverage, conditions for acoustic measurements and the aggregation (high patchiness leads to high variance) of the capelin.

The uncertainty is estimated by bootstrapping (see WKICE). The CV for the immature abundance was estimated to 0.15 in the 2024 autumn survey. The CV for the mature biomass was estimated to 0.25 in the 2024 autumn survey. In the two winter surveys in January (for the area east of Kolbeinsey Ridge) and February (for the western area) used for the assessment in 2025 the CV was 0.36 and 0.33, respectively.

While the autumn survey was affected by time constraints the distribution of immature and mature components of the capelin stock seemed to have been covered. The final estimate was based on the combination of the autumn survey and two winter surveys in January and February. The winter surveys had considerably lower SSB estimates than the autumn survey.

Although the final winter estimate of the fishable biomass derived from two different surveys conducted one month a part, it was not considered to add uncertainty to the assessment. It is because that considering the migration route of the stock, capelin in the east measured in January were for sure not measured again in the west in February. Furthermore, these were the most comprehensive measurements of capelin in these two areas of all the surveys conducted.

During the surveys, some components of the stock are likely to have been measured with the survey migration and others against it, and that can add bias and uncertainty to the estimates to unquantifiable degree.

Comparison with previous assessment and forecast

For the fishing season 2024/2025, the initial quota and intermediate advice were 0 t, but after the winter surveys a final TAC of 8 589 tonnes was given. The biomass estimates were roughly the same in autumn 2023 and 2024, but the immature component in autumn 2024 was higher than in the previous year.

Management considerations

The fishing season for capelin has since 1975 started in the period from late June to July/August, when surveys on the juvenile part of the stock the year before have resulted in the setting of an initial (preliminary) catch quota. During summer, the availability of plankton is at its highest and the fishable stock of capelin is feeding very actively over large areas between Iceland, Greenland and Jan Mayen, increasing rapidly in length, weight and fat content. By late September/beginning of October this period of rapid growth is over. The growth is fastest the in first two years, but the weight increase is highest in the year before spawning (Vilhjálmsson, 1994).

Given the large weight increase in the summer before spawning it is likely that there will be more biomass of maturing fish in autumn than in summer, even though the level of natural mortality is not well known during this time period. This should be considered for optimal timing of fishery in relation to yield and ecological impact. This is also supported by information for the Barents Sea capelin where it has been shown that fishing during autumn would maximize the yield, but from the ecosystem point of view a winter fishery were preferable (Gjøsæter et al., 2002). As the biology and role in the ecosystem of these two capelin stocks are similar, this is considered to be valid for the capelin in the Iceland-East Greenland-Jan Mayen area as well - until it is studied for this specific stock.

During the autumn surveys, juvenile and adult capelin are often found together. This should be considered during summer and autumn fishing because the survival rate of juvenile capelin that escape through the trawl net is unknown.

Ecosystem considerations

Capelin is an important forage fish and its dynamics are expected to have implications on the productivity of their predators.

The importance of capelin in East Greenlandic waters is not well documented but the effort has increased considerably during autumn surveys towards evaluation of the capelin role in the ecosystem e.g. by research on feeding of capelin, estimates of prey availability, predators’ distributions and environmental monitoring.

In Icelandic waters, capelin is the main single item in the diet of Icelandic cod, a key prey to several species of marine mammals and seabirds and also important as food for several other commercial fish species (see e.g. Vilhjálmsson, 2002, Singh et al., 2023).

Regulations and their effects

Over the years, the fishery has been closed during April–late June and the season has started in July/August or later, depending on the state of the stock.

Areas with high abundances of juvenile age 1 and 2 capelin (on the shelf region off NW-, N- and NE-Iceland) have usually been closed to the summer and autumn fishery.

It is permissible to transfer catches from the purse seine of one vessel to another vessel, in order to avoid slippage. However, if the catches are beyond the carrying capacity of the vessel and no other vessel is nearby, slippage is allowed. In recent years, reporting of such slippage has not been frequent. Industrial trawlers do not have the permission to slip capelin in order to harmonize catches to the processing.

In Icelandic waters, fishing with pelagic trawl is only allowed in limited area off the NE-coast (fishing in January) to protect juvenile capelin and to reduce the risk of affecting the spawning migration route (shuttering of migrating capelin schools by pelagic trawling has been hypothesized).

As a precautionary measure to protect juvenile capelin, the coastal states (Iceland, Greenland and Norway) have agreed that from 2021 fishing shall not start until 15. October.

References

Anon. 2015. Agreed Record of Conclusions of Coastal State consultations on the management of the capelin stock in the Iceland–East Greenland–Jan Mayen area. 2015. Reykjavík, Iceland. 7–8 May 2015. https://www.regjeringen.no/contentassets/37b66bdf33d84e99924bb27553641719/samledokument-lodde-mai-2015---agreed-records---bilateral-avtale.pdf Last accessed: 31 May 2025.

Anon. 2023. Framework arrangement between Greenland and Iceland on the conservation and management of capelin. 2023. Reykjávik, Iceland 3 July 2023. 4 pp. https://www.althingi.is/altext/pdf/154/fylgiskjol/s1655-f_I.pdf Last accessed: 31 May 2025.

Bardarson, B, Heilman, L, Jonsson, SÞ and Jansen, T 2024. Cruise report of acoustic assessment of the Iceland-East Greenland-Jan Mayen capelin stock in the autumn 2024. Kver Hafrannsóknastofnunar. KV2024-10. 18 pp.

Bardarson, B, Gudnason, K, Singh, W, Petursdottir, H, & Jonsson, SÞ (2021). Loðna (Mallotus villosus). Í Guðmundur J. Óskarsson (ritstj.), Staða umhverfis og vistkerfa í hafinu við Ísland og horfur næstu áratuga. Haf- og vatnarannsóknir, HV 2021-14, 31–34.

Christiansen, JS, Præbel, K, Siikavuopio, SI, Carscadden, JE 2008. Facultative semelparity in capelin Mallotus villosus (Osmeridae)-an experimental test of a life history phenomenon in a sub-arctic fish, Journal of Experimental Marine Biology and Ecology, Volume 360, Issue 1, 2008, Pages 47-55, ISSN 0022-0981, https://doi.org/10.1016/j.jembe.2008.04.003.

Engilbertsson, V, Óskarsson, GJ and Marteinsdóttir, G (2012). Inter-annual Variation in Fat Content of the Icelandic Capelin. ICES CM 2013/N:26.

Gjøsæter, H, Bogstad, B, and Tjelmeland, S 2002. Assessment methodology for Barents Sea capelin, Mallotus villosus (Müller). – ICES Journal of Marine Science, 59: 1086–1095.

Gudmundsdottir, A, and Vilhjálmsson, H 2002. Predicting Total Allowable Catches for Icelandic capelin, 1978–2001. ICES Journal of Marine Science, 59: 1105–1115.

Gudmundsdottir, A, and Sigurdsson, Th 2014. Growth of capelin in the Iceland-East Greenland-Jan Mayen area. NWWG 2014/WD:29.

ICES. 2015. Report of the Benchmark Workshop on Icelandic Stocks (WKICE), 26-30 January, 2015. ICES Headquarters. ICES CM 2015/ACOM:31. 335 pp.https://doi.org/10.17895/ices.pub.5295

ICES. 2023. Benchmark workshop on capelin (WKCAPELIN). ICES Scientific Reports. 5:62. 282 pp. https://doi.org/ices.pub.23260388

ICES 2024. Capelin (Mallotus villosus) in subareas 5 and 14 and Division 2.a west of 5°W (Iceland and Faroes grounds, East Greenland, Jan Mayen area). ICES Advice: Recurrent Advice. Report. https://doi.org/10.17895/ices.advice.25663980.v1

Jansen, T, Hansen, FT, Bardarson, B 2021. Larval drift dynamics, thermal conditions and the shift in juvenile capelin distribution and recruitment success around Iceland and East Greenland. Fisheries Research, 236.

MFRI. 2024. State of Marine Stocks and Advice 2024, advice on capelin. Marine and Freshwater Research Institute, 11 October 2024.

Singh, W, Ólafsdóttir, AH, Jónsson, SÞ, Óskarsson, GJ 2023. Capelin in a changing environment. Haf- og vatnarannsóknir, HV 2023-43. https://www.hafogvatn.is/static/research/files/capelin_2023_eng.pdf

Vilhjálmsson, H 1994. The Icelandic capelin stock. Capelin, Mallotus villosus (Müller), in the Iceland– Greenland–Jan Mayen area. Rit Fiskideildar, 13: 281 pp.

Vilhjálmsson, H 2002. Capelin (Mallotus villosus) in the Iceland–East Greenland–Jan Mayen ecosystem. ICES Journal of Marine Science: Journal du Conseil, 59: 870–883.

Vilhjálmsson, H 2007. Impact of changes in natural conditions on ocean resources. Law, science and ocean management 11, 225.